Goldstein Lab
UNC Chapel Hill


Our publications on tardigrades (more to come!)
Hypsibius dujardini water bear
Smith FW, Boothby TC, Giovannini I, Rebecchi L, Jockusch EL, and B Goldstein (2016). The compact body plan of tardigrades evolved by the loss of a large body region. Current Biology 26:224-229. (and a Dispatch article)

Boothby TC, Tenlen JR, Smith FW, Wang JR, Patanella KA, Osborne Nishimura E, Tintori SC, Li Q, Jones CD, Yandell M, Messina DN, Glasscock J, and Goldstein B (2016). Evidence for extensive horizontal gene transfer from the draft genome of a tardigrade. PNAS 112:15976-15981 and follow-up letter

Tenlen, J.R., S. McCaskill and B. Goldstein (2013). RNA interference can be used to disrupt gene function in tardigrades. Development Genes and Evolution 223:171-181. (and a F1000 review)

Gabriel WN, McNuff R, Patel SK, Gregory TR, Jeck WR, Jones CD and Goldstein B (2007).  The Tardigrade Hypsibius dujardini, a New Model for Studying the Evolution of Development. Developmental Biology 312:545-559. (and a F1000 review)

Gabriel WN and Goldstein B (2007). Segmental Expression of Pax3/7 and Engrailed Homologs in Tardigrade Development.  Development Genes and Evolution 217: 421-433. (and a F1000 review)

Goldstein B and Blaxter M (2002). Tardigrades. Current Biology 12: R475.


nsf water bears projectOur development of tardigrades as a model system has been supported by
National Science Foundation grants IOS 0235658, IOS 1257320, and IOS 1557432.

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